I used to suffer from them years ago.
Some possible causes:
Eye strain. I think this was almost certainly the cause of mine, early stages of needing glasses.
Bad posture, tension of back & neck muscles, especially when constantly balancing yourself boat fishing.
Dehydration, less likely perhaps if you normally take plenty to drink with you, but also worth weighing up if it's enough.
I would have included the sun, but you said you were wearing sunglasses all day.
I'm no doctor, but these things come to mind.